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Vladimir Bozukov

Personal information


assoc. prof. Vladimir Bozukov, PhD
Div. Palaeobotany and Palynology
Institute of Biodiversity and Ecosystem Research
Bulgarian Academy of Sciences
Acad. G. Bonchev Str., 23, BG-1113 Sofia, Bulgaria

Research interests

Palaeobotany, Palaeoecology, Palaeoclimatology, Biostratigraphy


1. Boev, Z., Bozukov, V. 2021. Satovcha (Blagoevgrad District, SW Bulgaria), a new middle Miocene avian locality. – C. R. Acad. Bulg. Sci., 74 (3): 406-411.
2. Bozukov, V. 1996a. Macclintockia basinervis (Rossm.) Knobloch in the fossil flora from Satovcha Graben in the Western Rhodopes. - Fitologija, 48: 68-72.
3. Bozukov, V. 1996b. The relationship between fossil flora from Satovcha Graben (the Western Rhodopes) and recent floras of North America and East Asia. - Phytologia Balcanica, 2(2): 32-37.
4. Bozukov, V. 1998a. Miocene macroflora of the Satovcha Graben (Western Rhodopes). I. Systematics. 1. Equisetophyta, Polypodiophyta, Pinophyta. - Phytologia Balcanica, 4(1-2): 3-12.
5. Bozukov, V. 1998b. Miocene macroflora of the Satovcha Graben (Western Rhodopes). I. Systematics. 2. Magnoliophyta: Lauraceae, Magnoliaceae, Cercidiphyllaceae, Hamamelidaceae, Ulmaceae, Moraceae. - Phytologia Balcanica, 4(3): 3-12.
6. Bozukov, V. 1999a. Miocene macroflora of the Satovcha Graben (Western Rhodopes). I. Systematics. 3. Fagaceae, Betulaceae. - Phytologia Balcanica, 5(1): 3-13.
7. Bozukov, V. 1999b. Miocene macroflora of the Satovcha Graben (Western Rhodopes). I. Systematics. 4. Magnoliophyta: Myricaceae, Juglandaceae, Theaceae, Salicaceae, Ericaceae, Myrsinaceae, Tiliaceae, Sterculiaceae, Fabaceae, Meliaceae, Anacardiaceae, Rutaceae, Aceraceae, Sapindaceae, Styracaceae, Cornaceae. - Phytologia Balcanica, 5(2-3): 47-62.
8. Bozukov, V. 2000. Miocene macroflora of the Satovcha Graben (Western Rhodopes). I. Systematics. 5. Magnoliophyta: Araliaceae, Aquifoliaceae, Celastraceae, Rhamnaceae, Vitaceae, Apocynaceae, Caprifoliaceae, Convolvulaceae, Macclintockia; Smilacaceae, Cyperaceae, Sparganiaceae, Typhaceae. - Phytologia Balcanica, 6(1): 15-30.
9. Bozukov, V. 2001. Miocene macroflora of the Satovcha Graben (Western Rhodopes). II. Paleofloristic, paleoecological and phytogeographical analysis. – Phytologia balcanica, 7(1): 13-33.
10. Bozukov, V. 2002. Miocene macroflara of the Satovcha Graben (Western Rhodopes). III. Comparative analysis of the Satovcha paleoflora with benchmark European macrofloras. Geological age of the paleoflora. – Phytologia Balcanica, 8(2): 165-180.
11. Bozukov, V. 2003. Tertiary history of fam. Betulaceae in Bulgaria. – Phytologia Balcanica, 9(2): 221-232.
12. Bozukov, V. 2005. Macclintockia basinervis (Rossm.) Knobl. In Cenozoic sediments in the Rhodopes Mt. Region (S Bulgaria). – Acta Palaeontol. Romaniae, 5: 11-15.
13. Bozukov, V. 2008. Genus Trichomanes in the European and Bulgarian Cenozoic macroflora. - Phytologia Balcanica, 14(1): 3-9.
14, Bozukov, V. 2011. Eugenia splendens Petrescu, G. Margarit & M. Margarit in the Bulgarian Paleoflora. - Acta Palaeontol. Romaniae, 7: 41-44.
15. Bozukov, V. 2016. Macroremains of edible plants from the Cenozoic of Bulgaria. – Phytol. Balcan., 22(2): 135-139.
16. Bozukov, V. 2018. Family Araliaceae in the Bulgarian macrofossil flora. Phytol. Balcan., 24(1): 3-10.
17. Bozukov, V. 2022. Revision of the species Bauhinia bulgarica Kitan. fil. (Fabaceae) known from the Bulgarian Upper Miocene-Lower Pliocene. – C. R. Acad. Bulg. Sci., 75(10): 1469-1474.
18. Bozukov, V., Djordjevic Milutinovic, D., Karamachoska, B., Garevska, B. 2022. Catalogue of the fossil Cenozoic plants of Republic of North Macedonia. – Acta Musei Macedonici Scientiarum Naturalium, 25(1): 7-58.
19. Bozukov, V. & Ivanov, D. 1995. Dawallia haidingeri Ett. a new species to Bulgarian flora - a leaf impression and spores in situ. - Phytologia Balcanica, 1(2): 15-18.
20. Bozukov, V. & Ivanov, D. 2020. New palaeobotanical data for the Late Oligocene Pernik Coal Basin (W Bulgaria ) - preliminary results. – C. R. Acad. Bulg. Sci., 73(11): 1562-1570.
21. Bozukov, V. & Ivanov, D. 2022. New data on the Middle Miocene flora of Northwest Bulgaria. – Phytol. Balcan., 28(2): 147-156.
22. Bozukov, V., Ivanov, D., Kováčová, M. 2013. A comparison between Badenian macroflora in Slovakia and Bulgaria. Proceedings of the National Conference with international participation GEOSCIENCES 2013, 65-67.
23. Bozukov, V., Ivanov, D., Todorov, O. 2023. New carpological data from the Satovcha paleoflora (SW Bulgaria). – Rev. Bulg. Geol. Soc., Geosciences 2023, 84(3): 145-148.
24. Бозуков, В., Иванов, Д. и Утешер, Т. 2010. Нови данни за разпространението на род Ficus (сем. Moraceae) в българската палеогенска флора. - БЪЛГАРСКО ГЕОЛОГИЧЕСКО ДРУЖЕСТВО, Национална конференция с международно участие „ГЕОНАУКИ 2010“, сборник разширени резюмета, 97-98 (English abstract).
25. Bozukov, V., Ivanov, D., Utescher, T. 2013. Ficus palamarevii sp. nov., a new subtropical element in the Bulgarian Paleogene flora. – C. R. Acad. Bulg. Sci., 12 (66): 1733-1738.
26. Bozukov, V. & Ivanova, R. 2015. New taxonomic data on the palaeoflora from the Satovcha Graben (SW Bulgaria). – C. R. Acad. Bulg. Sci., 68(5): 623-630.
27. Bozukov, V., Karamachoska, B. & Garevska, B. 2021. Specimens from the palaeobotanical collection of National Institution Museum of Natural History of Republic of North Macedonia, Skopje. – Acta Musei Macedonici Scientiarum Naturalium, 24(1): 7-18.
28. Bozukov, V., Kovacova, M., Durisova, A., Ivanov, D. 2018. First data about Araliaceae macroremains in Slovakia. – C. R. Acad. Bulg. Sci., 71(8) 1070-1075.
29. Bozukov, V., Kováčová, M., Ivanov, D. 2020. New data to the Cenozoic history of the genus Carpinus (Betulaceae) in Bulgaria. – C. R. Acad. Bulg. Sci., 73, 7: 971-977.
30. Bozukov, V. & Palamarev, E. 1992. Taxonomiche Zusammensetzung der Gattungen Populus L. und Alnus Gaerth. in der fossilen Flora von Satovcha Graben in West-Rhodopen (Bulgarien). - Documenta naturae, 76: 10-19.
31. Bozukov, V. & Palamarev, E. 1995. On the Tertiary History of the Theaceae in Bulgaria. - Flora mediterranea, 5: 171-184.
32. Bozukov, V., Palamarev, E. & Petkova, A. 2008. The fossil macroflora of the Vulche Pole Molasse formation (SE Bulgaria). - Phytologia Balcanica, 14(2): 173-184.
33. Bozukov, V., Todorov, O., Ivanova, R. 2023. New data on the Middle Miocene flora of the Satovcha Graben (SW Bulgaria). - Phytologia Balcanica, 29(3): 313-320.
34. Bozukov, V., Tsenov, B. 2012. Catalogue of the Cenozoic plants of Bulgaria (Eocene to Pliocene). Addendum and Corrigendum. – Phytologia Balcanica, 18(3): 237-261.
35. Bozukov, V., Tsenov, B. & Vatsev, M. 2009. New data about the fossil flora of Dospei Basin (South-west Bulgaria). – Dokl. Bulg. Akad. Nauk., 62(2):271-276.
36. Bozukov, V., Tsenov, B. & Vatsev, M. 2010. A first find of Botrychium (Ophioglossaceae) in Bulgarian palaeomacroflora. – Dokl. Bulg. Akad. Nauk., 63(6):889-892.
37. Bozukov, V., Tsenov, B., Valchev, V., Georgiev, V. & Tsoneva, S. 2012. Genus Ceratophyllum L. in the Bulgarian Neogene Flora. – Dokl. Bulg. Akad. Nauk., 65(7): 985-990.
38. Bozukov,V., Utescher, T. & Ivanov, D. 2009. Late Eocene to early Miocene climate and vegetation of Bulgaria. - Review of Palaeobotany & Palynology, 153: 360-374.
39. Bozukov, V., Utescher, T., Ivanov, D., Tsenov, B., Ashraf, A.R. & Mosbrugger, V. 2011. New results for the macroflora of the Beli Breg Lignite Basin, West Bulgaria. - Phytologia Balcanica, 17(1): 3-19.
40. Hristova, V., Ivanov, D., Bozukov, V. 2015. Palaeobotanical data for climate change during the late Miocene – early Pliocene in western Bulgaria. – C. R. Acad. Bulg. Sci., 68(7): 897-902.
41. Iamandei, S., Iamandei, E. & Bozukov, V. 2009. Is Pobitite kamani a petrified forest of Varna. In: Abstract book of SEVENTH ROMANIAN SYMPOSIUM ON PALEONTOLOGY, Cluj-Napoca, 22-24 October 2009 (Eds Bucur, I., Sasaran, E. & Pop, D.), p. 49.
42. Iamandei, S., Iamandei, E., Bozukov, V., Simov, N., Ivanov, D. 2023. First Evidence for Middle Miocene Petrified Wood Remains from the Bulgarian Black Sea Shore. – C. R. Acad. Bulg. Sci., 76(5): 742-750.
43. Iamandei, S., Iamandei, E., Bozukov, V. & Tzenov, B. 2011. Fossil woods from Rhodopes, Bulgaria. In: Abstract book of EIGHTH ROMANIAN SYMPOSIUM ON PALEONTOLOGY, BUCHAREST, 29-30 SEPTEMBER 2011 (Ed. Csiki Z.), p. 64.
44. Iamandei S., Iamandei E., Bozukov V., Tsenov B. 2016. Oligocene fossil woods from Rhodopes, Bulgaria. Acta Paleontologica Romaniae, 12(2):47-65.
45. Ivanov, D., Bozukov, V., Koleva-Rekalova, E. 2007. Late Miocene flora from SE Bulgaria: vegetation, landscape and climate reconstruction. - Phytologia Balcanica, 13(3): 281-292.
46. Ivanov, D., Kováčová, M., Bozukov, V., Kováč, M., Doláková, N. 2014. Late Miocene palaeoenvironmental dynamics in Central and Eastern Paratethys – preliminary results based on vegetation data. – C. R. Acad. Bulg. Sci., 67(4): 557-562.
47. Ivanov, D., Utescher, T., Ashraf, A.R., Mosbrugger, V., Bozukov, V. Djorgova, N., Slavomirova, E. 2012. Late Miocene Palaeoclimate and Ecosystem Dynamics in Southwestern Bulgaria - a Study Based on Pollen Data from the Gotse-Delchev Basin. - Turkish Journal of Earth Sciences (Turkish J. Earth Sci.), 21: 187-211.
48. Ivanov, D., Utesher, T., Ashraf, A. R., Mosbrugger, V., Slavomirova, E., Djorgova, N. & Bozukov, V. 2008. Vegetation structure and dynamics in the Late Miocene of Staniantsi Basin (West Bulgaria), first results. – Dokl. Bulg. Akad. Nauk., 61(2): 223-232.
49. Ivanov, D., Utescher, T., Djorgova, N., Bozukov, V., Ashraf, A.R. 2021. The late Miocene Beli Breg Basin (Bulgaria): palaeoecology and climate reconstructions based on pollen data. Palaeobio Palaeoenv. 101: 79-102.
50. Ivanov, M., Bozukov, V. 2017. Late Neogene braided rivers in the Blagoevgrad Graben: the first stages of the Late Miocene orthoplen destruction. – Rev. Bulg. Geol. Soc., 78(1–3), 39-55.
51. Ivanov, M., Bozukov, V. 2021. Stratigraphic relations between Boboshevo paleoflora and fossil floras with late Miocene age in Bulgaria. – C. R. Acad. Bulg. Sci., 74(11): 1642-1649.
52. Lazaridis, G., Kantiranis, N., Ioakeimidis, I., Bozukov, V. 2014. Excavations in the Early Villafranchian site Milia-5 (Grevena, Greece): a multidisciplinary approach. – Abstract book of VIth Internacional Conference on Mommoths and their Relatives. S.A.S.G., Aristotle University of Thessaloniki, Special Volume 102, 2.
53. Mantzouka, D., Bozukov, V., Argyrakos, G., Karakitsios, V. 2019. The first report of palaeobotanical remains from Zakynthos Island, Western Greece. – Phytol. Balcan., 25(2): 127-136.
54. Mantzouka, D., Ivanov, M., Bozukov, V. 2021. The first report of an ‘evergreen Castanopsis type’ wood (Fagaceae) for the Late Miocene–Early Pliocene of Europe (Bulgaria, Blagoevgrad Graben). – Geol. J., 56(2): 786-803.
55. Nel, A., Simov, N., Bozukov, V. & Marinov, M. 2016. New dragonflies and damselflies from Middle Miocene deposits in SW Bulgaria (Insecta: Odonata). – Palaeontologica Electronica, 19(3), 13 pp.
56. Palamarev, E. & Bozukov, V. 1992. On the Tertiary History of genus Acer Linnaeus in Bulgaria. - Geologica balcanica, 5: 61-70.
57. Palamarev, E. & Bozukov, V. 2004. The macroflora of Neogene sediments in the Elhovo Formation (SE Bulgaria). – Phytol. Balcan., 10(2-3): 131-146.
58. Palamarev, E., Bozukov, V. & Ivanov, D. 2002. Late Neogene floras from Bulgaria: vegetation and palaeoclimate estimates. - Acta Univ. Carolianae – Geologica, 46(4): 57-63.
59. Palamarev, E., Bozukov, V. & Staneva, K. 2001. Paleogene macroflora of the Borino-Teshel Graben in Western Rhodopes (Southwest Bulgaria). - Phytologia Balcanica, 7(3): 279-297.
60. Palamarev, E., Bozukov, V., Uzunova, K., Petkova, A. & Kitanov, G. 2005. Catalogue of the Cenozoic plants of Bulgaria (Eocene to Pliocene). - Phytologia Balcanica, 11(3): 215-364.
61. Palamarev, E., Ivanov, D. & Bozukov, V. 1999. Palaoflorenkomplexe im Zentralbalkanischen Raum und ihre Entwicklungsgeschichte von der Wende Oligozan/Miozan bis ins Villafranchien. - Flora Tertiaria Mediterranea, 6(5): 1-99.
62. Palamarev, E., Kitanov, G. & Bozukov, V. 1999. Paleogene flora from the central area of the Mesta Graben: the local flora of Boukovo (Western Rhodopes). - Phytologia Balcanica, 5(2-3): 27-46.
63. Palamarev, E., Kitanov, G., Bozukov, V. & Staneva, K. 1999. Fossil flora from Paleogene sediments in the northern area of the Mesta Graben in the Western Rhodopes. I. Systematics. - Phytologia Balcanica, 5(2-3): 3-25.
64. Palamarev, E., Kitanov, G., Staneva, K. & Bozukov, V. 2000. Fossil flora from Paleogene sediments in the northern area of the Mesta Graben in the Western Rhodopes. II. Analysis and stratigraphic importance of the flora. - Phytologia Balcanica, 6(1): 3-11.
65. Simov, N., Langourov, M., Pavlova, A., Hubenov, Z., Bozukov, V. 2021. First record of fossil Diptera (Insecta) in Miocene deposits in Bulgaria. – C. R. Acad. Bulg. Sci., 74 (2): 233-240.
66. Simov, N., Langourov, M., Sakalian, V., Bozukov, V. 2021. First fossil jewel beetle (Insecta: Coleoptera: Buprestidae) from Middle Miocene deposits in Bulgaria. - Historia naturalis bulgarica, 42: 31-34.
67. Tsenov, B. & Bozukov, V. 2011. Coexistence Aproach palaeoclimate reconstructions, based on the taxonomical composition of two macro floras from SW Bulgaria. – Anuarul Institutului Geologic al Romanei, 77: p. 33 (special issue).
68. Utescher, T., Ivanov, D., Harzhauser, M., Bozukov, V., Ashraf, A.R. & Mosbrugger, V. 2009. Cyclic climate and vegetation change in the late Miocene of Western Bulgaria. – Palaeogeography, Palaeoclimatology, Palaeoecology, 272: 99-114.


Plant and Fungal Diversity and Resources
Paleobotany and Palynology
Research group
Macro fossils


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